How do hummingbirds fly backwards? 

Wing Anatomy: Hummingbirds have highly specialized wings that allow them to generate lift in both the forward and backward directions. Their wings are long and narrow, with a high aspect ratio, enabling precise control of airflow and lift production. 

Rotary Joint: Hummingbirds have a unique shoulder joint that allows their wings to rotate in a full circle, rather than just up and down like most birds. This rotary joint enables them to produce lift on the upstroke as well as the downstroke, essential for sustained hovering and backward flight. 

Muscle Power: Hummingbirds have incredibly strong flight muscles relative to their body size, allowing them to generate the rapid wing beats necessary for sustained hovering and agile flight maneuvers, including flying backwards. 

Wingbeat Frequency: Hummingbirds beat their wings at incredibly high frequencies, often exceeding 50 beats per second during hovering and slow flight. This rapid wingbeat rate creates a continuous airflow over the wings, enabling precise control of flight direction and speed. 

Tail Control: Hummingbirds use their tail feathers to help control their flight, including backward flight. By adjusting the angle and position of their tail feathers, they can manipulate airflow and make precise adjustments to their flight path. 

Hummingbirds can fly vast distances and at high speeds. Hummingbirds can fly 30 mph in direct flight. These small insects also do aerial acrobatics and dives during courtship. Hummingbirds may dive over 45 mph.